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  • Drylie, T.P., Needham, H.R., Lohrer A.M., Hartland, A., Pilditch C.A. (2019).  Calcium carbonate alters the functional response of coastal sediments to eutrophication-induced acidification. Scientific Reports 9:12012. DOI: s41598-019-48549-8.

  • Drylie, T.P., Lohrer, A.M., Needham, H.R., Bulmer R.H., Pilditch C.A. (2018). Benthic primary production in emerged intertidal habitats provides resilience to high water column turbidity. Journal of Sea Research 142: 101-112.

  • Gladstone-Gallagher, R. V., Needham, H. R., Lohrer, A. M., Lundquist, C. J., & Pilditch, C. A. (2017). Site-dependent effects of bioturbator-detritus interactions alter soft-sediment ecosystem function. Marine Ecology Progress Series, 569: 145-161. doi:10.3354/meps12086

  • Needham, H. R., Pilditch, C. A., Lohrer, A. M., Thrush S. F. (2013) Density and habitat dependent effects of crab burrows on sediment erodibility. Journal of Sea Research 76: 94-104.

  • Widdicombe, S., Beesley, A., Berge, J.A., Dashfield, S.L., McNeill, C.L., Needham, H.R., Oxenvad, S. (2013) Impact of elevated levels of CO2 on animal mediated ecosystem function: The modification of sediment nutrient fluxes in burrowing urchins. Marine Pollution Bulletin 73: 416-427.

  • Needham, H. R., Pilditch, C., A. Lohrer, A. M., Thrush S. F. (2011) Context specific bioturbation mediates changes to ecosystem functioning. Ecosystems 14 (7): 1096-1109

  • Spicer, J. A., Widdicombe, S., Needham, H. R. Berge, J.A. (2011) Impact of CO2-acidified seawater on the extracellular acid-base balance of the northern sea urchin Strongylocentrotus dröebachiensis. Journal of Experimental Marine Biology and Ecology 407: 19-25.

  • Needham, H. R., Pilditch, C. A., Lohrer, A. M., Thrush S. F. (2010) Habitat dependence in the functional traits of a key burrowing species. Marine Ecology-Progress Series 414: 179- 193.

  • Widdicombe, S., Dashfield, S. L., McNeill, C. L., Needham, H. R., Beesley, A., McEvoy, A., Oxnevad, S. Clarke, K. R., Berge, J. A. (2009).  Effects of CO2 induced seawater acidification on infaunal diversity and sediment nutrient fluxes.  Marine Ecology-Progress Series  379 : 59-75.     

  • Widdicombe, S. & Needham, H. R. (2007). Impact of CO2 induced seawater acidification on the burrowing activity of Nereis virens (Sars 1835) and sediment nutrient flux. Marine Ecology-Progress Series 341: 111-122.

  • Clarke, K. R., Chapman, M. G., Somerfield, P. J., Needham, H. R. (2006). Dispersion-based weighting of species counts in assemblage analyses. Marine Ecology-Progress Series 320: 11-27.

  • Widdicombe, S., Austen, A.C., Kendall, M. A., Olsgard,F. Schanning, M.T., Dashfield, S. L., Needham, H. R. (2004) Importance of bioturbators for biodiversity maintenance: indirect effects of fishing disturbance. Marine Ecology-Progress Series 275:1-10.

Journal Articles

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